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[Objective]（1）To explore the distribution and characteristics of the clusters ofcalcium oxalate in the roots, stems, leaves, flowers, fruits and seeds of the Polygonaceaeplants.（2）To explore the law of the dynamic development about the clusters ofcalcium oxalate in the Polygonaceae plant organs of the different stages of growth；（3）Hope to provide some reference for the research of the identification of traditionalChinese medicine about the Clusters of calcium oxalate;（4）On the basis of previousstudies, preliminary exploration the functional role of calcium oxalate crystals in plants.[Methods] The topics combined the method of resources survey, give priority thePolygonaceae resources in Anhui, selected twenty kinds of Polygonaceae plants asrepresentatives, according to batches phenophase sampling. Producer the vegetativeorgans such as roots, rhizomes, stems, leaves and part of reproductive organs of thePolygonaceae plants.Observed clusters of calcium oxalate crystal morphology andstatistical comparisons.[Results]（1）Distribution law: In the Polygonaceae medicinal plants, thefollowing are distribution laws of Clusters of calcium oxalate in different organs.①In root, clusters of calcium oxalate distribution in parenchyma cells of the cortex,phloem ray and wood rays. The largest number of cortex containing clusters of calciumoxalate，and concentrated in the outer side of the cortex. In addition, it also found thatthe distribution of clusters of calcium oxalate in Rumex plant root and seasonal. have agreat relevance.②In stems, clusters of calcium oxalate distribution in the parenchyma cellsof thecortex and pith. The pith general relatively broad,the clusters of calcium oxalate isrelatively larger and more in Pith.Clusters of calcium oxalate centrally located in theouter side of the cortex, the clusters of calcium oxalate is scattered in Pith.③In leaves, the distribution that clusters of calcium oxalate on the leaf has a veryclose relationship with the level of veins, from the finest stockwork to the coarsest mainvein, as the veins roughness increases,clusters of calcium oxalate continue become increase and larger,and the growing trend is more pronounced. Horizontalcomparison,from the leaf margin to the the leaf middle of the main pulse, Clusters ofcalcium oxalate have the trend of increased and larger;longitudinal comparison,from thetip to the base of the leaf, clusters of calcium oxalate are growing larger. Plush, gillappendages on the leaf surface to the distribution of clusters of calcium oxalate in theleaves may also have an impact. Morphology of clusters calcium oxalate crystal inPolygonaceae genera of plants mature leaves are quite different, According to the pointof difference of clusters of calcium oxalate crystal,we can be listed in the Polygonaceaeeach genus belonging to retrieve table。As Polygonaceae leaf growth time increase,Clusters of calcium oxalate increasing,when the leaves withered,and there is no obviouschange in the number, size and distribution of Polygonaceae plant leaves in clusters ofcalcium oxalate. Clusters of calcium oxalate concentrated in the vascular bundlesaround Polygonaceae plant petiole.④In reproductive organs, perianth often containing clusters of calcium oxalate,however, the diameter is relatively small clusters of calcium oxalate，and clusters ofcalcium oxalate crystal of the perianth base are more,the clusters of calcium oxalate ofthe perianth edge relatively little or almost no, the stigma not contain clusters of calciumoxalate. The fruit calyx may exist clusters of calcium oxalate, But hard fruits or seedsgenerally do not contain clusters of calcium oxalate.（2）Morphological differences: In different groups of the Polygonaceae plants, morphology of Clusters of calcium oxalate crystal is significantly different.①In observed Polygonaceae plants, clusters of calcium oxalate maximum ofRumex plants is the biggest, edges and corners are very sharp, density is the least.②Clusters of calcium oxalate maximum of Antenoronv, Fagopyrum, Sect.Aconogonon, Homalocladium, and Sect. Bistorta is smaller range, the corners alsocompare blunt, density is relatively large.③Clusters of calcium oxalate maximum of Fallopia multiflora, Reynoutria Houtt.,Sect.Polygonum,Sect.Avicularia, Sect.Echinocaulon, Sect.Cephalophilon, Rheum is moderate range, edges and corners and some sharp and some blunt, density is moderaterange.④According to the morphological differences of clusters of calcium oxalate inPolygonaceae genera of plants, listed belong retrieval Form of Polygonaceae genera.（3）Dynamic development of the law:With the Polygonaceae medicinal plantgrowth and development,clusters of calcium oxalate is also increasing and increase, This phenomenon is common in Polygonaceae plant organs, the developmental changesclusters of calcium oxalate crystals in roots and leaves is the most obvious.Inaddition,also found that the Polygonaceae plant leaves stop growing and withered,thenumber, size and distribution of the calcium oxalate crystal clusters no significantchanges.（4）Defense function: In experiment, it found that some clues clusters of calciumoxalate in the Polygonaceae plant defense system have a closely related.①In the roots of Polygonaceae plants, the number of clusters of calcium oxalatecrystals change with the seasons.Polygonaceae plants Summer thrive root contain manyclusters of calcium oxalate,the winter hibernation root clusters of calcium oxalate less oralmost no. This phenomenon may be positively correlated with the activity frequency ofthe microorganisms and insects in soil.②Polygonaceae plants young leaves contain unusually large clusters of calciumoxalate, such clusters of calcium oxalate is greater than the clusters of calcium oxalatein the mature leaves, such as young leaves of Fagopyrum dibotrys (D. Don) Hara. Thisphenomenon may be related to the tender leaves are more vulnerable to abuse and needprotection.③Clusters of calcium oxalate often more and greater in important position the ofPolygonaceae plant organs. For example, in plant roots of Polygonaceae, Clusters ofcalcium oxalate centrally located in the outside of the cortex. This may be because ofthe roots contact with soil, the root bark is the first line of defense to protect the root, Therefore, the clusters of calcium oxalate in cortex is a largest number and concentrated in the outside of cortex. These clusters of calcium oxalate play the first defensive role.In Polygonaceae leaf, clusters of calcium oxalate concentrated in veins around, clusters of calcium oxalate around thicker veins is more and greater. In Polygonaceaestem and petiole, clusters of calcium oxalate only found in the stems of vascular bundlesscattered around. Because, vascular tissue is the most important organization in the plant, when the plant leaves and stems exposed to the air, vascular tissues need greaterprotection.In Polygonaceae reproductive organs, Clusters of calcium oxalate concentrated inthe base of perianth and fruit calyx, edge rarely. In addition, hard shell or seeds did notfind clusters of calcium oxalate, Probably because of the hard fruits and seeds have ahard shell protection, probably because of the hard fruits and seeds have hard protection,[Conclusion] The distribution of Clusters of calcium oxalate in Polygonaceaemedicinal plants have the certain regularity. Clusters of calcium oxalate of each speciesof Polygonaceae is differences, these differences are a certain value on the classificationof plants Polygonaceae. With the growth and development of medicinal plants ofPolygonaceae, clusters of calcium oxalate have a trend of increased and growing. Clusters of calcium oxalate may be an important part of the the Polygonaceae medicinalplant defense system. It does not need to produce clusters of calcium oxalate to defense.
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